<P> In vascular plants, the root is the organ of a plant that typically lies below the surface of the soil . Roots can also be aerial or aerating, that is growing up above the ground or especially above water . Furthermore, a stem normally occurring below ground is not exceptional either (see rhizome). Therefore, the root is best defined as the non-leaf, non-nodes bearing parts of the plant's body . However, important internal structural differences between stems and roots exist . </P> <P> The fossil record of roots--or rather, infilled voids where roots rotted after death--spans back to the late Silurian . Their identification is difficult, because casts and molds of roots are so similar in appearance to animal burrows . They can be discriminated using a range of features . </P> <P> The first root that comes from a plant is called the radicle . A root's four major functions are 1) absorption of water and inorganic nutrients, 2) anchoring of the plant body to the ground, and supporting it, 3) storage of food and nutrients, 4) vegetative reproduction and competition with other plants . In response to the concentration of nutrients, roots also synthesise cytokinin, which acts as a signal as to how fast the shoots can grow . Roots often function in storage of food and nutrients . The roots of most vascular plant species enter into symbiosis with certain fungi to form mycorrhizae, and a large range of other organisms including bacteria also closely associate with roots . </P> <P> When dissected, the arrangement of the cells in a root is root hair, epidermis, epiblem, cortex, endodermis, pericycle and, lastly, the vascular tissue in the centre of a root to transport the water absorbed by the root to other places of the plant . </P>

All of the following are functions of the root system of a plant except