<Tr> <Th> FMA </Th> <Td> 77826 </Td> </Tr> <Tr> <Td_colspan="2"> Anatomical terminology (edit on Wikidata) </Td> </Tr> <P> An otolith (Greek: ὠτο -, ōto - ear + λῐ́θος, líthos, a stone), also called statoconium or otoconium or statolith, is a calcium carbonate structure in the saccule or utricle of the inner ear, specifically in the vestibular system of vertebrates . The saccule and utricle, in turn, together make the otolith organs . These organs are what allows an organism, including humans, to perceive linear acceleration, both horizontally and vertically (gravity). They have been identified in both extinct and extant vertebrates . </P> <P> Counting the annual growth rings on the otoliths is a common technique in estimating the age of fish . </P>

In the otolith organs the otoliths are calcium carbonate particles that ____