<Li> Josh Brolin as Thanos: An intergalactic despot from Titan who longs to collect all six of the Infinity Stones in order to impose his will on all of reality, wanting to "re-balance the universe". Producer Kevin Feige added that Thanos believes the universe is becoming over-populated, which led to the destruction of his home moon Titan and is something he vowed not to let happen again, and also said "you could almost go so far as to say he is the main character of" the film . McFeely shared this sentiment, describing the film as his "hero journey" in addition to being the film's protagonist, stating, "Part of that is the things that (mean) the most to him . We wanted to show that . It wasn't just power; it wasn't just an ideal; it was people ." Brolin likened Thanos to "the Quasimodo of this time" and the novel Perfume, since Thanos was born deformed and considered a "freak" on Titan, while Joe Russo would reference The Godfather for Brolin at times, which Brolin felt helped "to emotionalize the whole thing". Brolin further added that he preferred playing Thanos over Cable in Deadpool 2 because of the amount of work that went into creating the character . Thanos does not wear armor for most of the film, which is symbolic of his growing power as he collects the Infinity Stones . In addition to voicing for the character, Brolin performed motion capture on set . </Li> <Li> Chris Pratt as Peter Quill / Star - Lord: The half - human, half - Celestial leader of the Guardians who was abducted from Earth as a child, and raised by a group of alien thieves and smugglers called the Ravagers . Pratt described his role in the film as "a cameo...you get to be a little more vibrant; a little more irreverent; a little bit more colorful if you want it to be". </Li> <P> Additionally, several other actors reprise their MCU roles: Danai Gurira as Okoye, the head of the Dora Milaje; Letitia Wright as T'Challa's sister Shuri; William Hurt as Thaddeus Ross, the U.S. Secretary of State; Kerry Condon as the voice of Stark's A.I. F.R.I.D.A.Y.; Winston Duke as M'Baku, the leader of Wakanda's mountain tribe the Jabari; Florence Kasumba as Ayo, a member of the Dora Milaje; Jacob Batalon as Parker's friend Ned; Isabella Amara as Parker's classmate Sally; Tiffany Espensen as Parker's classmate Cindy; and Ethan Dizon as Parker's classmate Tiny . Samuel L. Jackson and Cobie Smulders make uncredited cameos as Nick Fury and Maria Hill, the former director and deputy director of S.H.I.E.L.D, respectively, in the film's post-credits scene . </P> <P> Thanos' henchmen, known collectively in the comics as the Black Order and in the film as the "Children of Thanos", include Terry Notary as Cull Obsidian, Tom Vaughan - Lawlor as Ebony Maw, Carrie Coon as Proxima Midnight, and Michael James Shaw as Corvus Glaive . The foursome provided voices and motion - capture performances on set for their characters . As Coon was pregnant during filming, she mainly did facial capture for Proxima Midnight with some motion - capture, with stuntwoman Monique Ganderton standing - in and providing the rest on set . Ross Marquand portrays Johann Schmidt / Red Skull, the "Stonekeeper" and former Nazi commander of Hydra during World War II . Marquand replaces Hugo Weaving, who had expressed reluctance to reprise the character from Captain America: The First Avenger . Avengers co-creator Stan Lee makes a cameo appearance in the film as Parker's school bus driver, while screenwriter Stephen McFeely cameos as Secretary Ross's aide . David Cross was invited to make a cameo appearance as Tobias Fünke, his character from the sitcom Arrested Development, which the Russo brothers had previously worked on; this was prevented by a scheduling conflict, but Fünke still appears in the film as a specimen in the Collector's collection, played by an uncredited extra . </P>

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