<P> The Boveri--Sutton chromosome theory (also known as the chromosome theory of inheritance or the Sutton--Boveri theory) is a fundamental unifying theory of genetics which identifies chromosomes as the carriers of genetic material . It correctly explains the mechanism underlying the laws of Mendelian inheritance by identifying chromosomes with the paired factors (particles) required by Mendel's laws . It also states that chromosomes are linear structures with genes located at specific sites called loci along them . </P> <P> It states simply that chromosomes, which are seen in all dividing cells and pass from one generation to the next, are the basis for all genetic inheritance . Over a period of time random mutation creates changes in the DNA sequence of a gene . Genes are located on chromosomes . </P> <P> 2 . The process of synapsis (pseudo-reduction) consists in the union in pairs of the homologous members (i.e., those that correspond in size) of the two series . 3 . The first post-synaptic or maturation mitosis is equational and hence results in no chromosomic differentiation . 4 . The second post-synaptic division is a reducing division, resulting in the separation of the chromosomes which have conjugated in synapsis, and their relegation to different germ - cells . 5 . The chromosomes retain a morphological individuality throughout the various cell - divisions . </P> <P> W.S. Sutton, The Chromosomes in Heredity 1903 </P>

Who proposed that genes are located on chromosomes