<P> Oscar returns to the city with a new title of "Sharkslayer". Sykes becomes his manager and Oscar moves to the "top of the reef" to live in luxury . At the same time, Don Lino has everyone search for Lenny . When several sharks approach Oscar's neighborhood, his neighbors expect him to drive them away so he goes and runs into Lenny . Since he does not wish to return home, Lenny begs Oscar to let him stay at Oscar's house . Soon, Angie finds out about the lie and threatens to tell everyone . Oscar and Lenny stage an event in which Lenny pretends to terrorize the town and Oscar defeats him . Though this further cements Oscar's reputation, it greatly angers Don Lino . </P> <P> Oscar buys some Valentine's Day gifts for Angie, but before he can present them to her, he finds that Don Lino has kidnapped Angie to force a sit - down . Lenny attends disguised as a dolphin named Sebastian . Don Lino threatens to eat Angie if Oscar does not comply . Lenny grabs Angie into his mouth, but later regurgitates her . When Don Lino realizes that "Sebastian" is really Lenny, he chases Oscar through the reef . Oscar heads for the Whale Wash and ends up trapping both sharks . He is given an ovation by the other fish, but Oscar confesses that he is not a "Sharkslayer" and reveals the truth behind Frankie's death . He then tells Don Lino that everyone likes Lenny for who he is and urges him not to prejudge people before he knows them properly . Realizing that Oscar is right, Don Lino makes peace with him and Lenny, stating that he and his gang bear him no ill will . Oscar forsakes all the wealth he has acquired, makes peace with the sharks, becomes co-manager of the Whale Wash (now frequented by sharks, killer whales, and swordfish), and starts dating Angie . </P> <Ul> <Li> Will Smith as Oscar, an underachieving bluestreak cleaner wrasse worker in the Whale Wash of Reef City who wants to be rich, but his schemes always fail and he ends up owing five thousand clams to his boss . </Li> <Li> Robert De Niro as Don Edward Lino, a great white shark and leader of a mob of criminally - inclined great white sharks, who wants his two sons to take over his business and run it together, but becomes infuriated when Oscar gets in the way following one of his sons' death . James Gandolfini was originally set to voice the character, but had to drop out . </Li> <Li> Renée Zellweger as Angie, Oscar's angelfish best friend and co-worker, who harbors a secret unrequited love for him . </Li> <Li> Jack Black as Lenny Lino, Don Lino's youngest son, who is a vegetarian . </Li> <Li> Angelina Jolie as Lola, a seductive female gold - digger Siamese Fighting Fish whom Oscar develops a romantic interest in . </Li> <Li> Martin Scorsese as Sykes, the pufferfish owner of the Whale Wash and a loan shark to whom Oscar owes five thousand clams . He once worked with Don Lino, but was thrown out and called in his debts to pay off the gangster . </Li> <Li> Ziggy Marley and Doug E. Doug as Ernie and Bernie, two Jamaican jellyfish and Sykes' enforcers, who enjoy jabbing Oscar with their vicious stingers when he is in trouble with their boss . </Li> <Li> Michael Imperioli as Frankie Lino, Lenny's older brother and Don Lino's eldest and more savage son, who shares embarrassment of Lenny's vegetarian tendencies with his father . </Li> <Li> Vincent Pastore as Luca, Don Lino's green octopus "left - hand, right - hand man", with a tendency to state the obvious much to Lino's annoyance . He might be inspired to Luca Brasi from The Godfather . </Li> <Li> Peter Falk as Don Ira Feinberg, an elderly leopard shark and leader of a mob of criminally - inclined leopard sharks, who is a friend of Don Lino performing karaoke (badly) at the sharks' headquarters . </Li> <Li> Katie Couric as Katie Current, the local reporter of Reef City in the U.S. release . At the time, Katie Couric hosted Today in America . In the Australian release, then local Today co-host Tracy Grimshaw dubbed the lines . Fiona Phillips of the UK's GMTV performed the voice for the British release of the film . Cristina Parodi of Italy's Verissimo provided the Italian version of the character . </Li> <Li> David P. Smith as Crazy Joe, a deranged hermit crab who is Oscar's other friend . He normally lives in a dumpster near the Whale Wash . </Li> <Li> Bobb'e J. Thompson as Shortie #1 (voice) </Li> <Li> Phil LaMarr as Prawn Shop Owner (voice) </Li> <Li> Jenifer Lewis as Motown Turtle (voice) </Li> </Ul> <Li> Will Smith as Oscar, an underachieving bluestreak cleaner wrasse worker in the Whale Wash of Reef City who wants to be rich, but his schemes always fail and he ends up owing five thousand clams to his boss . </Li>

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