<Li> Bruce Willis as RJ, a raccoon who is duplicitous, selfish, and extremely intelligent . Despite his hard outer shell, he is revealed to have a sensitive personality, developing feelings of guilt over using his new - found companions to his own ends . </Li> <Li> Garry Shandling as Verne, a naturally cynical ornate box turtle, the leader of the foragers . He has his own ways of doing the daily tasks, but his world is turned upside - down when RJ introduces his free - spirited lifestyle into the mix . Though Verne genuinely cares for his family, he tends to condescend to them . A couple of running gags throughout the film are everyone calling him an amphibian and him correcting them with reptile (though Dwayne already knew he was a reptile, but first mistook him as an amphibian), as well as his shell constantly falling off . </Li> <Li> Steve Carell as Hammy, a hyperactive American red squirrel, whose mouth moves as fast as his feet and loves cookies . He is naive and childish in nature, with an extremely short attention span . The double entendre of Hammy's constantly trying to find his nuts is a third running gag in the film . </Li> <Li> Wanda Sykes as Stella, a short - tempered, sassy striped skunk who is constantly being told by the other foragers that she needs a man in her life . </Li>

Who plays the squirrel in over the hedge