<P> These social systems are affected by three main ecological factors: distribution of resources, group size, and predation . Within a social group there is a balance between cooperation and competition . Cooperative behaviors include social grooming (removing skin parasites and cleaning wounds), food sharing, and collective defense against predators or of a territory . Aggressive behaviors often signal competition for food, sleeping sites or mates . Aggression is also used in establishing dominance hierarchies . </P> <P> Several species of primates are known to associate in the wild . Some of these associations have been extensively studied . In the Tai Forest of Africa several species coordinate anti-predator behavior . These include the Diana monkey, Campbell's mona monkey, lesser spot - nosed monkey, western red colobus, king colobus and sooty mangabey, which coordinate anti-predator alarm calls . Among the predators of these monkeys is the common chimpanzee . </P> <P> The red - tailed monkey associates with several species, including the western red colobus, blue monkey, Wolf's mona monkey, mantled guereza, black crested mangabey and Allen's swamp monkey . Several of these species are preyed upon by the common chimpanzee . </P> <P> In South America, squirrel monkeys associate with capuchin monkeys . This may have more to do with foraging benefits to the squirrel monkeys than anti-predation benefits . </P>

Which of the following does the cladistic primate classification include