<Li> Marc Evan Jackson as Commander U.S. Central Command </Li> <Li> Katie Lowes as April the Residential Assistant </Li> <Ul> <Li> Peter Cullen as Optimus Prime The leader of the Autobots who transforms into a blue and red 1994 Peterbilt 379 semi-trailer truck . He is said to be the only Transformer capable of defeating The Fallen . </Li> <Li> Mark Ryan as Jetfire An ancient Decepticon - turned - Autobot Seeker who transforms into a Lockheed SR - 71 Blackbird . </Li> <Li> Reno Wilson as Mudflap An Autobot infiltrator and Skids' twin who transforms into a red 2007 Chevrolet Trax . </Li> <Li> Jess Harnell as Ironhide The Autobot weapons specialist and Optimus' new second - in - command who transforms into a black 2006 GMC Topkick C4500 . </Li> <Li> Robert Foxworth as Ratchet The Autobot medical officer who transforms into a yellow 2004 search and rescue Hummer H2 ambulance . </Li> <Li> André Sogliuzzo as Sideswipe The Autobot combat instructor who transforms into a silver 2009 Chevrolet Corvette Stingray . </Li> <Li> Grey DeLisle as Arcee A female Autobot who transforms into a pink Ducati 848 . </Li> <Li> Hugo Weaving as Megatron The Fallen's apprentice and the leader of the Decepticons who transforms into a Cybertronian hover tank . </Li> <Li> Tony Todd as The Fallen A rogue Prime who is the first and founder of the Decepticons, as well as the master of Megatron . </Li> <Li> Charlie Adler voices Starscream Megatron's second - in - command who transforms into a Lockheed Martin F - 22 Raptor . </Li> <Li> Frank Welker as <Ul> <Li> Soundwave The Decepticon communications officer who transforms into a Cybertronian satellite . </Li> <Li> Devastator A massive Decepticon who is the combination of nine Constructicons . </Li> <Li> Reedman, a razor - thin Decepticon composed of thousands of minuscule bead - like Decepticons . </Li> </Ul> </Li> <Li> Tom Kenny as <Ul> <Li> Wheelie A former Decepticon drone and later an Autobot who transforms into a blue radio - controlled toy monster truck . </Li> <Li> Skids An Autobot messenger and Mudflap's twin who transforms into a green 2007 Chevrolet Beat . </Li> </Ul> </Li> <Li> Calvin Wimmer voices Demolishor (credited as "Wheelbot") A huge Constructicon who transforms into a red and white Terex O&K RH 400 excavator . He hides out in Shanghai with the Decepticon Sideways . </Li> <Li> John DiCrosta as Scalpel (credited as "Doctor") A spider - like Decepticon who transforms into a microscope . </Li> <Li> Michael York as Prime #1 One of the Seven Primes . </Li> <Li> Kevin Michael Richardson as <Ul> <Li> Prime #2 One of the Seven Primes . </Li> <Li> Rampage (credited as "Skipjack") A Constructicon who transforms into a red Caterpillar D9 T bulldozer . He forms the left leg of Devastator . </Li> </Ul> </Li> <Li> Robin Atkin Downes as Prime #3 One of the Seven Primes </Li> </Ul> <Li> Peter Cullen as Optimus Prime The leader of the Autobots who transforms into a blue and red 1994 Peterbilt 379 semi-trailer truck . He is said to be the only Transformer capable of defeating The Fallen . </Li>

Voices of the transformers revenge of the fallen