<P> The series stars Clark Gregg, reprising his role of Phil Coulson from the films, as well as Ming - Na Wen, Brett Dalton, Chloe Bennet, Iain De Caestecker, and Elizabeth Henstridge . Nick Blood and Adrianne Palicki joined the cast for the second and third seasons, while Henry Simmons and Luke Mitchell had recurring roles in the second season before being promoted to the main cast for the third . John Hannah, who recurred in the third season, joined the main cast in the fourth . Additionally, several other characters from Marvel Cinematic Universe films and Marvel One - Shots also appear throughout the series, along with other characters based on various Marvel Comics properties . Several characters from the series also appear in the supplemental digital series Marvel's Agents of S.H.I.E.L.D.: Slingshot . </P> <P> This list includes the series' main cast, all guest stars deemed to have had recurring roles throughout the series, and any other guest who is otherwise notable . </P> <Table> <Tr> <Th> Character </Th> <Th> Portrayed by </Th> <Th_colspan="6"> Appearances </Th> </Tr> <Tr> <Th> First </Th> <Th> Season 1 </Th> <Th> Season 2 </Th> <Th> Season 3 </Th> <Th> Season 4 </Th> <Th> Season 5 </Th> </Tr> <Tr> <Th_colspan="8"> Main characters </Th> </Tr> <Tr> <Th> Phil Coulson </Th> <Td> Clark Gregg </Td> <Td> "Pilot" </Td> <Td_colspan="5"> Main </Td> </Tr> <Tr> <Th> Melinda May </Th> <Td> Ming - Na Wen </Td> <Td_colspan="5"> Main </Td> </Tr> <Tr> <Th> Grant Ward Hive </Th> <Td> Brett Dalton </Td> <Td_colspan="3"> Main </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Td> "4,722 Hours" </Td> <Td_colspan="2"> Does not appear </Td> <Td> Main </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Daisy "Skye" Johnson Quake </Th> <Td> Chloe Bennet </Td> <Td> "Pilot" </Td> <Td_colspan="5"> Main </Td> </Tr> <Tr> <Th> Leo Fitz </Th> <Td> Iain De Caestecker </Td> <Td_colspan="4"> Main </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Jemma Simmons </Th> <Td> Elizabeth Henstridge </Td> <Td_colspan="5"> Main </Td> </Tr> <Tr> <Th> Lance Hunter </Th> <Td> Nick Blood </Td> <Td> "Shadows" </Td> <Td> Does not appear </Td> <Td_colspan="2"> Main </Td> <Td> Does not appear </Td> <Td> Guest </Td> </Tr> <Tr> <Th> Bobbi Morse </Th> <Td> Adrianne Palicki </Td> <Td> "A Hen in the Wolf House" </Td> <Td> Does not appear </Td> <Td> Main </Td> <Td> Main </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Alphonso "Mack" MacKenzie </Th> <Td> Henry Simmons </Td> <Td> "Shadows" </Td> <Td> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="3"> Main </Td> </Tr> <Tr> <Th> Lincoln Campbell </Th> <Td> Luke Mitchell </Td> <Td> "Afterlife" </Td> <Td> Does not appear </Td> <Td> Recurring </Td> <Td> Main </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Holden Radcliffe </Th> <Td> John Hannah </Td> <Td> "The Singularity" </Td> <Td_colspan="2"> Does not appear </Td> <Td> Recurring </Td> <Td> Main </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th_colspan="8"> Recurring characters </Th> </Tr> <Tr> <Th> Mike Peterson Deathlok </Th> <Td> J. August Richards </Td> <Td> "Pilot" </Td> <Td> Recurring </Td> <Td> Guest </Td> <Td_colspan="3"> Does not appear </Td> </Tr> <Tr> <Th> Ian Quinn </Th> <Td> David Conrad </Td> <Td> "The Asset" </Td> <Td> Recurring </Td> <Td_colspan="4"> Does not appear </Td> </Tr> <Tr> <Th> Raina </Th> <Td> Ruth Negga </Td> <Td> "Girl in the Flower Dress" </Td> <Td_colspan="2"> Recurring </Td> <Td_colspan="3"> Does not appear </Td> </Tr> <Tr> <Th> Victoria Hand </Th> <Td> Saffron Burrows </Td> <Td> "The Hub" </Td> <Td> Recurring </Td> <Td_colspan="4"> Does not appear </Td> </Tr> <Tr> <Th> Davis </Th> <Td> Maximilian Osinski </Td> <Td> "Repairs" </Td> <Td> Guest </Td> <Td_colspan="2"> Does not appear </Td> <Td> Guest </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Anne Weaver </Th> <Td> Christine Adams </Td> <Td> "Seeds" </Td> <Td> Guest </Td> <Td> Recurring </Td> <Td_colspan="3"> Does not appear </Td> </Tr> <Tr> <Th> John Garrett The Clairvoyant </Th> <Td> Bill Paxton </Td> <Td> "T.A.H.I.T.I." </Td> <Td> Recurring </Td> <Td_colspan="4"> Does not appear </Td> </Tr> <Tr> <Th> Antoine Triplett </Th> <Td> B.J. Britt </Td> <Td_colspan="2"> Recurring </Td> <Td> Does not appear </Td> <Td> Guest </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Glenn Talbot </Th> <Td> Adrian Pasdar </Td> <Td> "Providence" </Td> <Td> Guest </Td> <Td_colspan="3"> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> The Koenigs </Th> <Td> Patton Oswalt </Td> <Td> Guest </Td> <Td> Recurring </Td> <Td> Does not appear </Td> <Td> Guest </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Calvin Johnson </Th> <Td> Kyle MacLachlan </Td> <Td> "Beginning of the End" </Td> <Td> Guest </Td> <Td> Recurring </Td> <Td_colspan="3"> Does not appear </Td> </Tr> <Tr> <Th> Daniel Whitehall </Th> <Td> Reed Diamond </Td> <Td> "Shadows" </Td> <Td> Does not appear </Td> <Td> Recurring </Td> <Td> Guest </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Sunil Bakshi </Th> <Td> Simon Kassianides </Td> <Td> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> <Td> Guest </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Kara Palamas Agent 33 </Th> <Td> Maya Stojan </Td> <Td> "Making Friends and Influencing People" </Td> <Td> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="3"> Does not appear </Td> </Tr> <Tr> <Th> Jiaying </Th> <Td> Dichen Lachman </Td> <Td> "The Things We Bury" </Td> <Td> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="3"> Does not appear </Td> </Tr> <Tr> <Th> Gordon </Th> <Td> Jamie Harris </Td> <Td> "What They Become" </Td> <Td> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="3"> Does not appear </Td> </Tr> <Tr> <Th> Andrew Garner Lash </Th> <Td> Blair Underwood </Td> <Td> "One of Us" </Td> <Td> Does not appear </Td> <Td> Guest </Td> <Td> Recurring </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Td> Matthew Willig </Td> <Td> "Laws of Nature" </Td> <Td_colspan="2"> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Robert Gonzales </Th> <Td> Edward James Olmos </Td> <Td> "Love in the Time of Hydra" </Td> <Td> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="3"> Does not appear </Td> </Tr> <Tr> <Th> Alisha Whitley </Th> <Td> Alicia Vela - Bailey </Td> <Td> "Scars" </Td> <Td> Does not appear </Td> <Td_colspan="2"> Guest </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Kebo </Th> <Td> Daz Crawford </Td> <Td> "S.O.S." </Td> <Td> Does not appear </Td> <Td_colspan="2"> Guest </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Rosalind Price </Th> <Td> Constance Zimmer </Td> <Td> "Laws of Nature" </Td> <Td_colspan="2"> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Luther Banks </Th> <Td> Andrew Howard </Td> <Td_colspan="2"> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Joey Gutierrez </Th> <Td> Juan Pablo Raba </Td> <Td_colspan="2"> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Werner von Strucker </Th> <Td> Spencer Treat Clark </Td> <Td> "Purpose in the Machine" </Td> <Td_colspan="2"> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Gideon Malick </Th> <Td> Powers Boothe </Td> <Td> "Among Us Hide ..." </Td> <Td_colspan="2"> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Giyera </Th> <Td> Mark Dacascos </Td> <Td> "Many Heads, One Tale" </Td> <Td_colspan="2"> Does not appear </Td> <Td> Recurring </Td> <Td_colspan="2"> Does not appear </Td> </Tr> <Tr> <Th> Elena "Yo - Yo" Rodriguez </Th> <Td> Natalia Cordova - Buckley </Td> <Td> "Bouncing Back" </Td> <Td_colspan="2"> Does not appear </Td> <Td_colspan="2"> Recurring </Td> <Td> Guest </Td> </Tr> <Tr> <Th> J.T. James Hellfire </Th> <Td> Axle Whitehead </Td> <Td> "Paradise Lost" </Td> <Td_colspan="2"> Does not appear </Td> <Td> Recurring </Td> <Td> Guest </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Anderson </Th> <Td> Alexander Wraith </Td> <Td_colspan="2"> Does not appear </Td> <Td_colspan="2"> Guest </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Piper </Th> <Td> Briana Venskus </Td> <Td> "Failed Experiments" </Td> <Td_colspan="2"> Does not appear </Td> <Td_colspan="2"> Guest </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Aida "Ophelia" Madame Hydra </Th> <Td> Mallory Jansen </Td> <Td> "Ascension" </Td> <Td_colspan="2"> Does not appear </Td> <Td> Guest </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Robbie Reyes Ghost Rider </Th> <Td> Gabriel Luna </Td> <Td> "The Ghost" </Td> <Td_colspan="3"> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Gabe Reyes </Th> <Td> Lorenzo James Henrie </Td> <Td_colspan="3"> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Lucy Bauer </Th> <Td> Lilli Birdsell </Td> <Td_colspan="3"> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Jeffrey Mace Patriot </Th> <Td> Jason O'Mara </Td> <Td> "Meet the New Boss" </Td> <Td_colspan="3"> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Ellen Nadeer </Th> <Td> Parminder Nagra </Td> <Td> "Uprising" </Td> <Td_colspan="3"> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Burrows </Th> <Td> Patrick Cavanaugh </Td> <Td_colspan="3"> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Eli Morrow </Th> <Td> José Zúñiga </Td> <Td> "Let Me Stand Next to Your Fire" </Td> <Td_colspan="3"> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Anton Ivanov The Superior </Th> <Td> Zach McGowan </Td> <Td> "Hot Potato Soup" </Td> <Td_colspan="3"> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> <Tr> <Th> Hope MacKenzie </Th> <Td> Jordan Rivera </Td> <Td> "Identity and Change" </Td> <Td_colspan="3"> Does not appear </Td> <Td> Recurring </Td> <Td> Does not appear </Td> </Tr> </Table> <Tr> <Th> Character </Th> <Th> Portrayed by </Th> <Th_colspan="6"> Appearances </Th> </Tr>

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