<P> Over periods of ecological changes, biogeography includes the study of plant and animal species in: their past and / or present living refugium habitat; their interim living sites; and / or their survival locales . As writer David Quammen put it, "...biogeography does more than ask Which species? and Where . It also asks Why? and, what is sometimes more crucial, Why not? ." </P> <P> Modern biogeography often employs the use of Geographic Information Systems (GIS), to understand the factors affecting organism distribution, and to predict future trends in organism distribution . Often mathematical models and GIS are employed to solve ecological problems that have a spatial aspect to them . </P> <P> Biogeography is most keenly observed on the world's islands . These habitats are often much more manageable areas of study because they are more condensed than larger ecosystems on the mainland . Islands are also ideal locations because they allow scientists to look at habitats that new invasive species have only recently colonized and can observe how they disperse throughout the island and change it . They can then apply their understanding to similar but more complex mainland habitats . Islands are very diverse in their biomes, ranging from the tropical to arctic climates . This diversity in habitat allows for a wide range of species study in different parts of the world . </P> <P> One scientist who recognized the importance of these geographic locations was Charles Darwin, who remarked in his journal "The Zoology of Archipelagoes will be well worth examination". Two chapters in On the Origin of Species were devoted to geographical distribution . </P>

What is pangaea and how does it relate to the concept of biogeography