<Li> Angelina Jolie as Lola, a seductive female gold - digger lionfish whom Oscar develops a romantic interest in . </Li> <Li> Martin Scorsese as Sykes, the pufferfish owner of the Whale Wash and a loan shark to whom Oscar owes five thousand clams . He once worked with Don Lino, but was thrown out and called in his debts to pay off the gangster . </Li> <Li> Ziggy Marley and Doug E. Doug as Ernie and Bernie, two Jamaican jellyfish and Sykes' enforcers, who enjoy jabbing Oscar with their vicious stingers when he is in trouble with their boss . </Li> <Li> Michael Imperioli as Frankie Lino, Lenny's older brother and Don Lino's eldest and more savage son, who shares embarrassment of Lenny's vegetarian tendencies with his father . </Li>

Who plays ernie and bernie in shark tale