<P> Water potential is the potential energy of water per unit volume relative to pure water in reference conditions . Water potential quantifies the tendency of water to move from one area to another due to osmosis, gravity, mechanical pressure, or matrix effects such as capillary action (which is caused by surface tension). The concept of water potential has proved useful in understanding and computing water movement within plants, animals, and soil . Water potential is typically expressed in potential energy per unit volume and very often is represented by the Greek letter ψ . </P> <P> Water potential integrates a variety of different potential drivers of water movement, which may operate in the same or different directions . Within complex biological systems, many potential factors may be operating simultaneously . For example, the addition of solutes lowers the potential (negative vector), while an increase in pressure increases the potential (positive vector). If flow is not restricted, water will move from an area of higher water potential to an area that is lower potential . A common example is water with a dissolved salt, such as sea water or the fluid in a living cell . These solutions have negative water potential, relative to the pure water reference . With no restriction on flow, water will move from the locus of greater potential (pure water) to the locus of lesser (the solution); flow proceeds until the difference in potential is equalized or balanced by another water potential factor, such as pressure or elevation . </P>

Why water potential of a cell is normally a negative value