<P> The animals are soon able to regroup, initially believing themselves to be at the San Diego Zoo . Upon exploring, however, they come across a pack of lemurs, led by King Julien XIII the ring - tailed lemur, and learn their true location . Alex blames Marty for their predicament and attempts to signal for help to get back to civilization . Marty, on the other hand, finds the wild to be exactly what he was looking for, with Gloria and Melman soon joining him in enjoying the island . Alex eventually comes around, but, deprived from the raw steaks he was provided with at the zoo, his prey drive begins to show as hunger kicks in . The group is accepted by the lemurs, though King Julien's adviser, Maurice the aye - aye, cautions them about Alex's predatory nature . King Julien ignores Maurice's concerns and persuades the group to help the lemurs fend off the fossa, who hunt the lemurs as prey . While Alex initially scares the fossa away and is worshiped by the lemurs, later, compelled by hunger, he attacks Marty . Realizing that Alex is now a threat, King Julien banishes him to the far side of the island where the fossa live . Seeing what has happened to Alex, and how difficult it is to survive with so many predators around the island, Marty begins to regret his decision to leave the zoo . </P> <P> The penguins, having been to Antarctica and found that it wasn't what they had in mind, land the ship at Madagascar . Seeing this as a chance to return Alex to New York, Marty rushes after his friend against the wishes of Melman and Gloria . Marty attempts to convince the now grizzled, starving Alex to return, but Alex refuses out of fear of attacking Marty again . The penguins, Gloria, and Melman go to find Marty, but are trapped by the fossa . At the last minute, Alex overcomes his predatory instincts and scares the fossa away from the lemur territory forever . The lemurs regain their respect for Alex, and the penguins help him satisfy his hunger through sushi . As the lemurs throw a farewell celebration for the foursome, the penguins decide not to break the news that the ship has run out of fuel . </P> <Ul> <Li> Ben Stiller as Alex, a lion . Tom McGrath explained that "Ben Stiller was the first actor we asked to perform, and we knew we wanted his character, Alex, to be a big performing lion with a vulnerable side ." </Li> <Li> Chris Rock as Marty, a zebra . McGrath explained the character: "Marty is a guy who thinks there might be more to life than what's in the zoo . We wanted his character to be energetic, so we listened to Chris Rock ." </Li> <Li> David Schwimmer as Melman, a hypochondriac giraffe who is afraid of germs . When they were looking for a voice actor for Melman, they listened to Schwimmer's voice on Friends and, according to McGrath, thought that it "sounded really neat ." </Li> <Li> Jada Pinkett Smith as Gloria, a strong, confident, but sweet hippopotamus . McGrath said that they found all these traits in Pinkett Smith's voice, when they listened to her . </Li> <Li> Sacha Baron Cohen as King Julien XIII, a ring - tailed lemur and the king of the lemurs . King Julien was initially only meant to be a "two - line" character until auditioning Baron Cohen improvised eight minutes of dialogue in an Indian accent . </Li> <Li> Cedric the Entertainer as Maurice, an aye - aye and King Julien's royal advisor . </Li> <Li> Andy Richter as Mort, a Goodman's mouse lemur . </Li> <Li> Tom McGrath as Skipper, the leader of penguins . McGrath, who was also the film's co-director and co-writer, initially only lent his voice to the temporary tracks . Growing up with films starring tough actors like John Wayne, Charlton Heston, and Robert Stack, McGrath wanted Stack for the voice of Skipper . Stack was approached about voicing the character, but died two weeks before production on the animation began . After that, DreamWorks Animation CEO Jeffrey Katzenberg decided to keep the temporary voice, with McGrath explaining: "People were used to me doing that voice . We knew it worked when we screened it ." Many character's traits were based on Stack's work . McGrath especially emphasized The Untouchables, a 1959 television crime drama series starring Stack . </Li> <Li> Chris Miller as Kowalski, a penguin and Skipper's right hand . </Li> <Li> Jeffrey Katzenberg as Rico, a smart and silent penguin who is only expressed through grunts and squeals . Mireille Soria, the film's producer, commented on Katzenberg's uncredited role: "The irony for us is that he's the one who doesn't talk . There's something very Dadaistic about that, isn't there?" </Li> <Li> Christopher Knights as Private, an eager, lowly penguin . Knights was also an assistant editor on the film . </Li> <Li> Conrad Vernon as Mason, a chimpanzee (Phil, the other chimpanzee, is unvoiced) </Li> <Li> Eric Darnell and Tom McGrath as the fossa </Li> <Li> Eric Darnell and David P. Smith as lemurs </Li> <Li> Elisa Gabrielli as Nana, an elderly New Yorker </Li> <Li> Bob Saget as an unspecified off - screen zoo animal </Li> <Li> David Cowgill as a police horse </Li> <Li> Stephen Apostolina as a police officer </Li> </Ul> <Li> Ben Stiller as Alex, a lion . Tom McGrath explained that "Ben Stiller was the first actor we asked to perform, and we knew we wanted his character, Alex, to be a big performing lion with a vulnerable side ." </Li>

Who does the voice of alex the lion in madagascar