<P> A 2009 genetic clustering study, which genotyped 1327 polymorphic markers in various African populations, identified six ancestral clusters through Bayesian analysis and fourteen ancestral clusters through STRUCTURE analysis within the continent . The clustering corresponded closely with ethnicity, culture and language . </P> <P> In addition, whole genome sequencing analysis of modern populations inhabiting Sub-Saharan Africa has observed several primary inferred ancestry components: a Pygmy - related component carried by the Mbuti and Biaka Pygmies in Central Africa, a Khoisan - related component carried by Khoisan - speaking populations in Southern Africa, a Niger - Congo - related component carried by Niger - Congo - speaking populations throughout Sub-Saharan Africa, a Nilo - Saharan - related component carried by Nilo - Saharan - speaking populations in the Nile Valley and African Great Lakes, and a West Eurasian - related component carried by Afroasiatic - speaking populations in the Horn of Africa and Nile Valley . </P> <P> Sub-Saharan Africa has several large cities . Lagos is a city in the Nigerian state of Lagos . The city, with its adjoining conurbation, is the most populous in Nigeria, and the second most populous on the African continent after Cairo, Egypt . It is one of the fastest growing cities in the world, and also one of the most populous urban agglomerations . Lagos is a major financial centre in Africa; the megacity has the highest GDP, and also houses one of the largest and busiest ports on the continent . </P> <P> Dar es Salaam is the former capital as well as the most populous city in Tanzania and a regionally important economic centre . It is located on the Swahili coast . </P>

Explain how when and by whom the borders of the countries in subsaharan africa were created