<Li> 1.5, as for FADH, if hydrogen atoms (2H + 2e) are transferred from cytosolic NADH + H to mitochondrial FAD by the glycerol phosphate shuttle located in the inner mitochondrial membrane . </Li> <Li> 2.5 in case of malate - aspartate shuttle transferring hydrogen atoms from cytosolic NADH + H to mitochondrial NAD </Li> <P> So finally we have, per molecule of glucose </P> <Ul> <Li> Substrate - level phosphorylation: 2 ATP from glycolysis + 2 ATP (directly GTP) from Krebs cycle </Li> <Li> Oxidative phosphorylation <Ul> <Li> 2 NADH + H from glycolysis: 2 × 1.5 ATP (if glycerol phosphate shuttle transfers hydrogen atoms) or 2 × 2.5 ATP (malate - aspartate shuttle) </Li> <Li> 2 NADH + H from the oxidative decarboxylation of pyruvate and 6 from Krebs cycle: 8 × 2.5 ATP </Li> <Li> 2 FADH from the Krebs cycle: 2 × 1.5 ATP </Li> </Ul> </Li> </Ul>

How many net atp molecules are produced by the citric acid cycle